{
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},
"headers": [
["alpha;", "Synonymous substitution rate at a site"],
["β-", "Non-synonymous substitution rate at a site for the negative/neutral evolution component"],
["p-", "Mixture distribution weight allocated to β-; loosely -- the proportion of the tree evolving neutrally or under negative selection"],
["β+", "Non-synonymous substitution rate at a site for the positive/neutral evolution component"],
["p+", "Mixture distribution weight allocated to β+; loosely -- the proportion of the tree evolving neutrally or under positive selection"],
["LRT", "Likelihood ratio test statistic for episodic diversification, i.e., p+ > 0 and β+ > α"],
["p-value", "Asymptotic p-value for episodic diversification, i.e., p+ > 0 and β+ > α"],
["# branches under selection", "The (very approximate and rough) estimate of how many branches may have been under selection at this site, i.e., had an empirical Bayes factor of 100 or more for the β+ rate"],
["Total branch length", "The total length of branches contributing to inference at this site, and used to scale dN-dS"],
["MEME LogL", "Site Log-likelihood under the MEME model"],
["FEL LogL", "Site Log-likelihood under the FEL model"]
]
},
"analysis":{
"authors":"Sergei L. Kosakovsky Pond, Steven Weaver",
"citation":"Detecting Individual Sites Subject to Episodic Diversifying Selection. _PLoS Genet_ 8(7): e1002764.",
"contact":"spond@temple.edu",
"info":"MEME (Mixed Effects Model of Evolution)\n estimates a site-wise synonymous (α) and a two-category mixture of non-synonymous\n (β-, with proportion p-, and β+ with proportion [1-p-]) rates, and\n uses a likelihood ratio test to determine if β+ > α at a site.\n The estimates aggregate information over a proportion of branches at a site,\n so the signal is derived from\n episodic diversification, which is a combination of strength of selection [effect size] and\n the proportion of the tree affected. A subset of branches can be selected\n for testing as well, in which case an additional (nuisance) parameter will be\n inferred -- the non-synonymous rate on branches NOT selected for testing. Multiple partitions within a NEXUS file are also supported\n for recombination - aware analysis.\n ",
"requirements":"in-frame codon alignment and a phylogenetic tree",
"version":"2.1.2"
},
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"original name":"epi_isl_704837_England_2020_11_22_null_1"
},
"epi_isl_706698_Wales_2020_11_06_null_1":{
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"original name":"epi_isl_706698_Wales_2020_11_06_null_1"
},
"epi_isl_706883_England_2020_null_2":{
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"Nucleotide GTR":0,
"original name":"epi_isl_706883_England_2020_null_2"
},
"epi_isl_707958_Saarland_2020_03_25_null_1":{
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"original name":"epi_isl_707958_Saarland_2020_03_25_null_1"
},
"epi_isl_708786_Western_Australia_2020_03_27_null_1":{
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"Nucleotide GTR":0,
"original name":"epi_isl_708786_Western_Australia_2020_03_27_null_1"
},
"epi_isl_708846_England_2020_12_03_null_2":{
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"original name":"epi_isl_708846_England_2020_12_03_null_2"
},
"epi_isl_712422_Hovedstaden_2020_11_30_null_1":{
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"Nucleotide GTR":0,
"original name":"epi_isl_712422_Hovedstaden_2020_11_30_null_1"
},
"epi_isl_713508_Hovedstaden_2020_11_23_null_2":{
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"Nucleotide GTR":0.001686113588749801,
"original name":"epi_isl_713508_Hovedstaden_2020_11_23_null_2"
}
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"EBF site 102 (partition 1)":{
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"EBF site 103 (partition 1)":{
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"EBF site 13 (partition 1)":{
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"EBF site 26 (partition 1)":{
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"EBF site 27 (partition 1)":{
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"EBF site 29 (partition 1)":{
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"EBF site 3 (partition 1)":{
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"EBF site 31 (partition 1)":{
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"EBF site 33 (partition 1)":{
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},
"EBF site 34 (partition 1)":{
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"EBF site 36 (partition 1)":{
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},
"EBF site 37 (partition 1)":{
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},
"EBF site 4 (partition 1)":{
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},
"EBF site 41 (partition 1)":{
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},
"EBF site 48 (partition 1)":{
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"EBF site 49 (partition 1)":{
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"EBF site 50 (partition 1)":{
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},
"EBF site 51 (partition 1)":{
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},
"EBF site 53 (partition 1)":{
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},
"EBF site 54 (partition 1)":{
"attribute type":"branch label",
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},
"EBF site 61 (partition 1)":{
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},
"EBF site 63 (partition 1)":{
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},
"EBF site 68 (partition 1)":{
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},
"EBF site 7 (partition 1)":{
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},
"EBF site 71 (partition 1)":{
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},
"EBF site 74 (partition 1)":{
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},
"EBF site 75 (partition 1)":{
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},
"EBF site 8 (partition 1)":{
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},
"EBF site 89 (partition 1)":{
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},
"EBF site 9 (partition 1)":{
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},
"EBF site 92 (partition 1)":{
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"EBF site 93 (partition 1)":{
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"EBF site 95 (partition 1)":{
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},
"Global MG94xREV":{
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"display order":1
},
"Nucleotide GTR":{
"attribute type":"branch length",
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},
"original name":{
"attribute type":"node label",
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}
}
},
"data partitions":{
"0":{
"coverage": [
[0, 1, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15, 16, 17, 18, 19, 20, 21, 22, 23, 24, 25, 26, 27, 28, 29, 30, 31, 32, 33, 34, 35, 36, 37, 38, 39, 40, 41, 42, 43, 44, 45, 46, 47, 48, 49, 50, 51, 52, 53, 54, 55, 56, 57, 58, 59, 60, 61, 62, 63, 64, 65, 66, 67, 68, 69, 70, 71, 72, 73, 74, 75, 76, 77, 78, 79, 80, 81, 82, 83, 84, 85, 86, 87, 88, 89, 90, 91, 92, 93, 94, 95, 96, 97, 98, 99, 100, 101, 102, 103, 104, 105, 106, 107, 108, 109, 110, 111, 112, 113, 114, 115, 116, 117, 118, 119, 120, 121, 122, 123, 124, 125, 126, 127, 128, 129, 130, 131, 132, 133, 134, 135, 136, 137, 138, 139, 140, 141, 142, 143, 144, 145, 146, 147, 148, 149, 150, 151, 152, 153, 154, 155, 156, 157, 158, 159, 160, 161, 162, 163, 164, 165, 166, 167, 168, 169, 170, 171, 172, 173, 174, 175, 176, 177, 178, 179, 180, 181, 182, 183, 184, 185, 186, 187, 188, 189, 190, 191, 192, 193, 194, 195, 196, 197]
],
"name":"meme.filter.default"
}
},
"fits":{
"Global MG94xREV":{
"AIC-c":5382.818378625388,
"Equilibrium frequencies": [
[0.03822960114694726],
[0.01351933692398675],
[0.02403787782864348],
[0.05193150930367305],
[0.02911052729638136],
[0.01029451039894302],
[0.01830401776853703],
[0.0395440594139595],
[0.0103437444419216],
[0.003657913291546855],
[0.006503904245034899],
[0.01405105584689999],
[0.02823116412447807],
[0.00998353652940644],
[0.01775109480154166],
[0.03834952284091316],
[0.01525310815750509],
[0.005394037660154572],
[0.009590797167562675],
[0.02071998933880218],
[0.01161471760238769],
[0.004107374281517386],
[0.00730306240097105],
[0.01577755972157445],
[0.004127018017262807],
[0.001459459303597651],
[0.002594972270682336],
[0.005606186518559166],
[0.011263863259984],
[0.003983299796723079],
[0.007082453407800259],
[0.01530095533648721],
[0.03598474580469079],
[0.01272547680494126],
[0.02262636536601744],
[0.04888207319672321],
[0.02740114711043603],
[0.009690013203794347],
[0.01722919954287686],
[0.03722201862968041],
[0.009736356206817353],
[0.003443119363604041],
[0.006121992748394631],
[0.01322597300962387],
[0.02657342044676213],
[0.009397299826948971],
[0.01670874440288144],
[0.03609762565549655],
[0.008264129496846549],
[0.03174480525674235],
[0.01779474604793255],
[0.006292850568177904],
[0.01118891954555733],
[0.0241725781127722],
[0.002236017143410683],
[0.003975720645048336],
[0.008589159789351673],
[0.01725720702747418],
[0.006102757789039748],
[0.01085092759915328],
[0.02344237921441801]
],
"Log Likelihood":-1904.313418267846,
"Rate Distributions":{
"non-synonymous/synonymous rate ratio for *background*": [
[0.7907130744508751, 1]
],
"non-synonymous/synonymous rate ratio for *test*": [
[0.4206399721350306, 1]
]
},
"display order":1,
"estimated parameters":779
},
"Nucleotide GTR":{
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"Equilibrium frequencies": [
[0.3252445297300179],
[0.1796505068272878],
[0.2035038156410188],
[0.2916011478016755]
],
"Log Likelihood":-1964.582685198452,
"Rate Distributions":{
"Substitution rate from nucleotide A to nucleotide C":0.125239360201612,
"Substitution rate from nucleotide A to nucleotide G":1,
"Substitution rate from nucleotide A to nucleotide T":0.02541668730675812,
"Substitution rate from nucleotide C to nucleotide G":0,
"Substitution rate from nucleotide C to nucleotide T":5.296852327218998,
"Substitution rate from nucleotide G to nucleotide T":0.5732006751686097
},
"display order":0,
"estimated parameters":771
}
},
"input":{
"file name":"/data/shares/veg/SARS-CoV-2/SARS-CoV-2-devel/data/fasta/2020-12-22/sequences.nsp8.compressed.filtered.fas",
"number of sequences":383,
"number of sites":198,
"partition count":1,
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