{
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},
"headers": [
["alpha;", "Synonymous substitution rate at a site"],
["β-", "Non-synonymous substitution rate at a site for the negative/neutral evolution component"],
["p-", "Mixture distribution weight allocated to β-; loosely -- the proportion of the tree evolving neutrally or under negative selection"],
["β+", "Non-synonymous substitution rate at a site for the positive/neutral evolution component"],
["p+", "Mixture distribution weight allocated to β+; loosely -- the proportion of the tree evolving neutrally or under positive selection"],
["LRT", "Likelihood ratio test statistic for episodic diversification, i.e., p+ > 0 and β+ > α"],
["p-value", "Asymptotic p-value for episodic diversification, i.e., p+ > 0 and β+ > α"],
["# branches under selection", "The (very approximate and rough) estimate of how many branches may have been under selection at this site, i.e., had an empirical Bayes factor of 100 or more for the β+ rate"],
["Total branch length", "The total length of branches contributing to inference at this site, and used to scale dN-dS"],
["MEME LogL", "Site Log-likelihood under the MEME model"],
["FEL LogL", "Site Log-likelihood under the FEL model"]
]
},
"analysis":{
"authors":"Sergei L. Kosakovsky Pond, Steven Weaver",
"citation":"Detecting Individual Sites Subject to Episodic Diversifying Selection. _PLoS Genet_ 8(7): e1002764.",
"contact":"spond@temple.edu",
"info":"MEME (Mixed Effects Model of Evolution)\n estimates a site-wise synonymous (α) and a two-category mixture of non-synonymous\n (β-, with proportion p-, and β+ with proportion [1-p-]) rates, and\n uses a likelihood ratio test to determine if β+ > α at a site.\n The estimates aggregate information over a proportion of branches at a site,\n so the signal is derived from\n episodic diversification, which is a combination of strength of selection [effect size] and\n the proportion of the tree affected. A subset of branches can be selected\n for testing as well, in which case an additional (nuisance) parameter will be\n inferred -- the non-synonymous rate on branches NOT selected for testing. Multiple partitions within a NEXUS file are also supported\n for recombination - aware analysis.\n ",
"requirements":"in-frame codon alignment and a phylogenetic tree",
"version":"2.1.2"
},
"branch attributes":{
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"Node841":{
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"EBF site 100 (partition 1)":{
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"EBF site 115 (partition 1)":{
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"EBF site 120 (partition 1)":{
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"EBF site 121 (partition 1)":{
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"EBF site 122 (partition 1)":{
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"EBF site 123 (partition 1)":{
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"EBF site 153 (partition 1)":{
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"EBF site 157 (partition 1)":{
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"EBF site 158 (partition 1)":{
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"EBF site 160 (partition 1)":{
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"EBF site 161 (partition 1)":{
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"EBF site 164 (partition 1)":{
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"EBF site 165 (partition 1)":{
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"EBF site 166 (partition 1)":{
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"EBF site 168 (partition 1)":{
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"EBF site 169 (partition 1)":{
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"EBF site 170 (partition 1)":{
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"EBF site 172 (partition 1)":{
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"EBF site 173 (partition 1)":{
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"EBF site 176 (partition 1)":{
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"EBF site 177 (partition 1)":{
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"EBF site 18 (partition 1)":{
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"EBF site 180 (partition 1)":{
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"EBF site 184 (partition 1)":{
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"EBF site 185 (partition 1)":{
"attribute type":"branch label",
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"EBF site 186 (partition 1)":{
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"EBF site 187 (partition 1)":{
"attribute type":"branch label",
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"EBF site 188 (partition 1)":{
"attribute type":"branch label",
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"EBF site 19 (partition 1)":{
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"EBF site 190 (partition 1)":{
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"EBF site 192 (partition 1)":{
"attribute type":"branch label",
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"EBF site 193 (partition 1)":{
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"EBF site 195 (partition 1)":{
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"EBF site 196 (partition 1)":{
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"EBF site 197 (partition 1)":{
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"EBF site 198 (partition 1)":{
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"EBF site 2 (partition 1)":{
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"EBF site 20 (partition 1)":{
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"EBF site 201 (partition 1)":{
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"EBF site 202 (partition 1)":{
"attribute type":"branch label",
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"EBF site 203 (partition 1)":{
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"EBF site 204 (partition 1)":{
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"EBF site 205 (partition 1)":{
"attribute type":"branch label",
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"EBF site 206 (partition 1)":{
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"EBF site 207 (partition 1)":{
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"EBF site 208 (partition 1)":{
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"EBF site 209 (partition 1)":{
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"EBF site 21 (partition 1)":{
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"EBF site 210 (partition 1)":{
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"EBF site 211 (partition 1)":{
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"EBF site 212 (partition 1)":{
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"EBF site 213 (partition 1)":{
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"EBF site 214 (partition 1)":{
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"EBF site 215 (partition 1)":{
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"EBF site 217 (partition 1)":{
"attribute type":"branch label",
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"attribute type":"branch label",
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"EBF site 219 (partition 1)":{
"attribute type":"branch label",
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"EBF site 64 (partition 1)":{
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"EBF site 65 (partition 1)":{
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},
"EBF site 68 (partition 1)":{
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},
"EBF site 7 (partition 1)":{
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},
"EBF site 70 (partition 1)":{
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},
"EBF site 71 (partition 1)":{
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},
"EBF site 73 (partition 1)":{
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},
"EBF site 74 (partition 1)":{
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},
"EBF site 76 (partition 1)":{
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},
"EBF site 77 (partition 1)":{
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},
"EBF site 78 (partition 1)":{
"attribute type":"branch label",
"display order":80
},
"EBF site 79 (partition 1)":{
"attribute type":"branch label",
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},
"EBF site 8 (partition 1)":{
"attribute type":"branch label",
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},
"EBF site 80 (partition 1)":{
"attribute type":"branch label",
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},
"EBF site 82 (partition 1)":{
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},
"EBF site 85 (partition 1)":{
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},
"EBF site 87 (partition 1)":{
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},
"EBF site 89 (partition 1)":{
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},
"EBF site 9 (partition 1)":{
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},
"EBF site 90 (partition 1)":{
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},
"EBF site 91 (partition 1)":{
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},
"EBF site 93 (partition 1)":{
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},
"EBF site 95 (partition 1)":{
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"EBF site 96 (partition 1)":{
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},
"EBF site 98 (partition 1)":{
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},
"Global MG94xREV":{
"attribute type":"branch length",
"display order":1
},
"Nucleotide GTR":{
"attribute type":"branch length",
"display order":0
},
"original name":{
"attribute type":"node label",
"display order":-1
}
}
},
"data partitions":{
"0":{
"coverage": [
[0, 1, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12, 13, 14, 15, 16, 17, 18, 19, 20, 21, 22, 23, 24, 25, 26, 27, 28, 29, 30, 31, 32, 33, 34, 35, 36, 37, 38, 39, 40, 41, 42, 43, 44, 45, 46, 47, 48, 49, 50, 51, 52, 53, 54, 55, 56, 57, 58, 59, 60, 61, 62, 63, 64, 65, 66, 67, 68, 69, 70, 71, 72, 73, 74, 75, 76, 77, 78, 79, 80, 81, 82, 83, 84, 85, 86, 87, 88, 89, 90, 91, 92, 93, 94, 95, 96, 97, 98, 99, 100, 101, 102, 103, 104, 105, 106, 107, 108, 109, 110, 111, 112, 113, 114, 115, 116, 117, 118, 119, 120, 121, 122, 123, 124, 125, 126, 127, 128, 129, 130, 131, 132, 133, 134, 135, 136, 137, 138, 139, 140, 141, 142, 143, 144, 145, 146, 147, 148, 149, 150, 151, 152, 153, 154, 155, 156, 157, 158, 159, 160, 161, 162, 163, 164, 165, 166, 167, 168, 169, 170, 171, 172, 173, 174, 175, 176, 177, 178, 179, 180, 181, 182, 183, 184, 185, 186, 187, 188, 189, 190, 191, 192, 193, 194, 195, 196, 197, 198, 199, 200, 201, 202, 203, 204, 205, 206, 207, 208, 209, 210, 211, 212, 213, 214, 215, 216, 217, 218, 219, 220, 221, 222, 223, 224, 225, 226, 227, 228, 229, 230, 231, 232, 233, 234, 235, 236, 237, 238, 239, 240, 241, 242, 243, 244, 245, 246, 247, 248, 249, 250, 251, 252, 253, 254, 255, 256, 257, 258, 259, 260, 261, 262, 263, 264, 265, 266, 267, 268, 269, 270, 271, 272, 273, 274, 275, 276, 277, 278, 279, 280, 281, 282, 283, 284, 285, 286, 287, 288, 289, 290, 291, 292, 293, 294, 295, 296, 297, 298, 299, 300, 301, 302, 303, 304, 305, 306, 307, 308, 309, 310, 311, 312, 313, 314, 315, 316, 317, 318, 319, 320, 321, 322, 323, 324, 325, 326, 327, 328, 329, 330, 331, 332, 333, 334, 335, 336, 337, 338, 339, 340, 341, 342, 343, 344, 345, 346, 347, 348, 349, 350, 351, 352, 353, 354, 355, 356, 357, 358, 359, 360, 361, 362, 363, 364, 365, 366, 367, 368, 369, 370, 371, 372, 373, 374, 375, 376, 377, 378, 379, 380, 381, 382, 383, 384, 385, 386, 387, 388, 389, 390, 391, 392, 393, 394, 395, 396, 397, 398, 399, 400, 401, 402, 403, 404, 405, 406, 407, 408, 409, 410, 411, 412, 413, 414, 415, 416, 417, 418, 419, 420, 421, 422, 423, 424, 425, 426, 427, 428, 429, 430, 431, 432, 433, 434, 435, 436, 437, 438, 439, 440, 441, 442, 443, 444, 445, 446, 447, 448, 449, 450, 451, 452, 453, 454, 455, 456, 457, 458, 459, 460, 461, 462, 463, 464, 465, 466, 467, 468, 469, 470, 471, 472, 473, 474, 475, 476, 477, 478, 479, 480, 481, 482, 483, 484, 485, 486, 487, 488, 489, 490, 491, 492, 493, 494, 495, 496, 497, 498, 499, 500, 501, 502, 503, 504, 505, 506, 507, 508, 509, 510, 511, 512, 513, 514, 515, 516, 517, 518, 519, 520, 521, 522, 523, 524, 525, 526, 527, 528, 529, 530, 531, 532, 533, 534, 535, 536, 537, 538, 539, 540, 541, 542, 543, 544, 545, 546, 547, 548, 549, 550, 551, 552, 553, 554, 555, 556, 557, 558, 559, 560, 561, 562, 563, 564, 565, 566, 567, 568, 569, 570, 571, 572, 573, 574, 575, 576, 577, 578, 579, 580, 581, 582, 583, 584, 585, 586, 587, 588, 589, 590, 591, 592, 593, 594, 595, 596, 597, 598, 599, 600]
],
"name":"meme.filter.default"
}
},
"fits":{
"Global MG94xREV":{
"AIC-c":21962.168931416,
"Equilibrium frequencies": [
[0.03290059496205524],
[0.01229753969476754],
[0.01087616970394493],
[0.0395784851084966],
[0.02472010329287806],
[0.009239846630540432],
[0.00817188986468087],
[0.02973758502501421],
[0.01801514956855218],
[0.00673367813505964],
[0.005955388471713039],
[0.02167171535191413],
[0.02623142921489242],
[0.009804747980778563],
[0.008671498981925008],
[0.03155566735961993],
[0.01846980192297316],
[0.006903617474523253],
[0.006105685941065302],
[0.02221864927392214],
[0.01387742111841654],
[0.005187083615404384],
[0.004587551906312977],
[0.01669414506676978],
[0.01011338076188725],
[0.003780165723782332],
[0.003343247913107846],
[0.01216611098803991],
[0.01472585229277232],
[0.005504209067298237],
[0.004868023473622113],
[0.01771478377067627],
[0.03380796238245404],
[0.01263669425665198],
[0.01117612422025292],
[0.04067002244931391],
[0.0254018604581427],
[0.009494672897715951],
[0.008397262890148964],
[0.03055771961054853],
[0.0185119904254111],
[0.006919386635658235],
[0.006119632476457733],
[0.02226940084900507],
[0.02695486732558744],
[0.0100751536951202],
[0.00891065075627118],
[0.03242594294351389],
[0.01223876519587908],
[0.03938932486289021],
[0.02460195675954114],
[0.009195685979792761],
[0.00813283337503419],
[0.02959545808731935],
[0.006701495392178743],
[0.005926925463517994],
[0.02156813819408604],
[0.0261060594949728],
[0.009757887457161539],
[0.00863005467518361],
[0.03140485113278214]
],
"Log Likelihood":-9605.222334852699,
"Rate Distributions":{
"non-synonymous/synonymous rate ratio for *background*": [
[0.4695025249380118, 1]
],
"non-synonymous/synonymous rate ratio for *test*": [
[0.9192447802560163, 1]
]
},
"display order":1,
"estimated parameters":1373
},
"Nucleotide GTR":{
"AIC-c":23911.56081206418,
"Equilibrium frequencies": [
[0.3033216461160401],
[0.1867508644284476],
[0.1946577791999161],
[0.3152697102555962]
],
"Log Likelihood":-9750.327301817608,
"Rate Distributions":{
"Substitution rate from nucleotide A to nucleotide C":0.2396998084257031,
"Substitution rate from nucleotide A to nucleotide G":1,
"Substitution rate from nucleotide A to nucleotide T":0.1360253542952463,
"Substitution rate from nucleotide C to nucleotide G":0.036028038572919,
"Substitution rate from nucleotide C to nucleotide T":2.266384736369818,
"Substitution rate from nucleotide G to nucleotide T":0.8090289298741782
},
"display order":0,
"estimated parameters":2203
}
},
"input":{
"file name":"/data/shares/veg/SARS-CoV-2/SARS-CoV-2/data/fasta/2020-06-11/sequences.helicase.compressed.fas",
"number of sequences":1099,
"number of sites":601,
"partition count":1,
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ales_2020_03_25_null_1,epi_isl_425134_North_Rhine_Westphalia_2020_03_11_null_1)Node1349,(epi_isl_452986_England_2020_05_14_null_3,epi_isl_453588_England_2020_04_27_null_7,epi_isl_457402_England_2020_04_29_null_1)Node1352,(epi_isl_443228_Singapore_2020_03_18_null_3,epi_isl_438599_England_2020_04_22_null_3,epi_isl_461000_Limburg_2020_05_08_null_16)Node1359,(epi_isl_433580_Scotland_2020_04_02_null_1,epi_isl_463893_Tuzla_2020_05_25_null_1)Node1365,epi_isl_457798_Maryland_2020_03_20_null_16,(epi_isl_456314_Southern_2020_04_03_null_1,epi_isl_461457_Washington_2020_05_29_null_2)Node1381,(epi_isl_448437_England_2020_04_19_null_1,epi_isl_456282_Southern_2020_03_29_null_1,epi_isl_463993_Toronto_2020_03_13_null_123)Node1373,(epi_isl_443702_England_2020_03_27_null_1,epi_isl_462054_England_2020_04_19_null_1)Node1378,((epi_isl_444177_England_2020_04_11_null_1,epi_isl_458875_England_2020_04_01_null_6)Node1392,epi_isl_426788_Victoria_2020_03_23_null_1,epi_isl_456357_Canterbury_2020_04_06_null_41)Node1388,epi_isl_423339_England_2020_03_27_null_1,epi_isl_452158_Utah_2020_04_10_null_1,((epi_isl_443935_England_2020_03_29_null_2,epi_isl_427082_Victoria_2020_03_28_null_1,epi_isl_462368_Singapore_2020_04_01_null_4)Node1397,epi_isl_440722_England_2020_04_05_null_2)Node1396,(epi_isl_438732_England_2020_04_26_null_1,epi_isl_442991_England_2020_04_10_null_3)Node1404,epi_isl_438381_England_2020_03_31_null_1,(epi_isl_446625_Wales_2020_04_09_null_2,(epi_isl_453313_England_2020_04_04_null_1,epi_isl_453342_England_2020_04_12_null_1)Node1425,(epi_isl_423870_England_2020_03_18_null_1,epi_isl_423888_England_2020_03_18_null_2)Node1421,epi_isl_456154_Tumaco_2020_04_26_null_20)Node1418,(epi_isl_447182_Michigan_2020_03_21_null_1,epi_isl_462181_Voroux_les_Liers_2020_03_17_null_18)Node1410,((epi_isl_430357_New_York_2020_04_03_null_1,epi_isl_458804_England_2020_04_14_null_11)Node1414,epi_isl_448978_England_2020_04_21_null_3)Node1413,epi_isl_444848_Unknown_2020_03_31_null_4,(epi_isl_445784_Wales_2020_04_09_null_4,epi_isl_440152_England_2020_03_21_null_3,epi_isl_442351_England_2020_04_10_null_1)Node940,(epi_isl_416396_Shanghai_2020_02_10_null_1,epi_isl_461368_Zuid_Holland_2020_05_02_null_8)Node945,(epi_isl_438998_Scotland_2020_04_22_null_1,epi_isl_452467_Andalusia_2020_03_10_null_1,epi_isl_459671_Scotland_2020_03_24_null_64)Node949,(epi_isl_429103_Guangzhou_2020_02_09_null_1,epi_isl_456784_Scotland_2020_05_07_null_2)Node954,(epi_isl_430479_Victoria_2020_03_31_null_1,epi_isl_456423_Victoria_2020_04_16_null_2)Node958,(epi_isl_448503_England_2020_04_06_null_3,epi_isl_448536_England_2020_04_04_null_1)Node961,((epi_isl_438905_Scotland_2020_04_14_null_1,epi_isl_457343_England_2020_05_13_null_3)Node980,epi_isl_453405_England_2020_04_09_null_1)Node979,((epi_isl_459510_Scotland_2020_03_27_null_3,epi_isl_450471_California_2020_03_21_null_2,epi_isl_462731_Michigan_2020_04_23_null_128)Node968,epi_isl_456523_Victoria_2020_05_07_null_5)Node967,((epi_isl_451693_Basel_Land_2020_03_14_null_1,epi_isl_461645_Scotland_2020_04_24_null_12)Node975,epi_isl_452667_Basque_Country_2020_03_26_null_2)Node974,(epi_isl_438691_England_2020_04_24_null_1,epi_isl_446833_Wales_2020_04_15_null_1)Node985,(epi_isl_449105_England_2020_04_08_null_1,epi_isl_453335_England_2020_04_12_null_1)Node988,(epi_isl_452654_Basque_Country_2020_03_27_null_1,epi_isl_454072_Portugal_2020_03_19_null_4)Node1005,(epi_isl_458121_Muscat_2020_04_20_null_1,epi_isl_453319_England_2020_04_10_null_1,epi_isl_458124_Muscat_2020_05_03_null_30)Node1008,(epi_isl_447943_England_2020_05_02_null_1,epi_isl_456113_New_Jersey_2020_04_26_null_1)Node995,epi_isl_460329_Massachusetts_2020_03_27_null_34,(epi_isl_451331_Sichuan_2020_02_02_null_3,epi_isl_451393_Sichuan_2020_02_15_null_3)Node1000,epi_isl_454672_California_2020_05_12_null_1)"
}
},
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